Trees or shrubs (or climbers).
Leaves opposite; simple; ±serrate margin, teeth widely distanced; no stipule; brochidodromous venation ("boxes" along the margin).
3-many ±tepals; 8-many stamens, Monoecious or dioecious.
Fruit and seed
Apocarpous superior ovaries; fruit achene (one-seeded "soft nut"), ovaries often (partly) enclosed in the receptacle.
- Vascular plants