Ursus

UrsusLinnaeus, 1758

Hyväksytty nimi, Maailman nisäkkäiden luettelo 2005

The close relationship of the four species included herein has been generally recognized by morphological and molecular studies (Goldman et al., 1989; Hendey, 1980a; Kurtén and Anderson, 1980; Shields and Kocher, 1991). Allen (1938) proposed a close relationship between thibetanus and americanus. Thenius (1953), Goldman et al. (1989), and Shields and Kocher (1991) gave support to the monophyly of arctos with maritimus. Synonyms allocated according to McKenna and Bell (1997).

• Wilson, D and Reeder, D. (eds.) 2005. Mammal Species of the World. A Taxonomic and Geographic Reference. 3rd ed. -- 2142 pp. Johns Hopkins University Press.